''Baccharis pilularis'', called coyote brush (or bush), chaparral broom, and bush baccharis, is a

shrub A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...

in the family

Asteraceae The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae w ...

native to

California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...

Oregon Oregon () is a U.S. state, state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington (state), Washington, while the Snake River delineates much of it ...

Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...

, and

Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...

. There are reports of isolated populations in

New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...

, most likely introduced.

Distribution and habitat

The plants are found in a variety of habitats, from coastal bluffs, oak woodlands, and grasslands, including on hillsides and in canyons, below . Coyote brush is known as a secondary pioneer plant in communities such as

coastal sage scrub Coastal sage scrub, also known as coastal scrub, CSS, or soft chaparral, is a low scrubland plant community of the California coastal sage and chaparral subecoregion, found in coastal California and northwestern coastal Baja California. It is w ...

and

chaparral Chaparral ( ) is a shrubland plant community and geographical feature found primarily in the U.S. state of California, in southern Oregon, and in the northern portion of the Baja California Peninsula in Mexico. It is shaped by a Mediterranean c ...

. It does not regenerate under a closed shrub canopy because seedling growth is poor in the shade.

Coast live oak ''Quercus agrifolia'', the California live oak, or coast live oak, is a highly variable, often evergreen oak tree, a type of live oak, native to the California Floristic Province. It may be shrubby, depending on age and growing location, but is ...

California bay ''Umbellularia californica'' is a large hardwood tree native to coastal forests and the Sierra foothills of California, and to coastal forests extending into Oregon. It is endemic to the California Floristic Province. It is the sole species in th ...

, '' Rhus integrifolia'', and other shade producing species replace coastal sage scrub and other coyote bush-dominated areas, particularly when there has not been a wildfire or heavy grazing. In California

grassland A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur natur ...

s, it comes in late and invades and increases in the absence of fire or grazing. Coyote bush invasion of grasslands is important because it helps the establishment of other coastal sage species. However, establishment of coyote bush can be concerning because it also displaces highly biodiverse grassland habitat that are important to carbon storage and resilient to wildfires. After grassland restoration, coyote bush can be a major concern and plant invader that overtakes grassland habitat, especially if restoration activities are limited and nonperiodic.

Description

The ''Baccharis pilularis'' shrub is generally smaller than in height. Erect plants are generally mixed (and intergrade completely) with prostrate plants. It is glabrous and generally sticky. The stems are prostrate to erect which branches spreading or ascending. The leaves are long and are

entire Entire may refer to: * Entire function, a function that is holomorphic on the whole complex plane * Entire (animal), an indication that an animal is not neutered * Entire (botany) This glossary of botanical terms is a list of definitions of ...

to toothed and oblanceolate to obovate, with three principal veins. The

flower heads A pseudanthium (Greek for "false flower"; ) is an inflorescence that resembles a flower. The word is sometimes used for other structures that are neither a true flower nor a true inflorescence. Examples of pseudanthia include flower heads, compos ...

are in a leafy

panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...

. The involucres are hemispheric to bell shaped. This species is

dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproductio ...

(

pistillate Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...

and

staminate The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filam ...

flowers occur on separate plants). Both

and

heads are long.

Phyllaries In botanical terminology, a phyllary, also known an involucral bract or tegule, is a single bract of the involucre of a composite flower. The involucre is the grouping of bracts together. Phyllaries are reduced leaf-like structures that form one or ...

are in 4–6 series, ovate, and glabrous. The receptacles are convex to conic and honeycombed. The

flowers range from 20–30 and there are 19–43

flowers. This and other ''

Baccharis ''Baccharis'' is a genus of perennials and shrubs in the aster family (Asteraceae). They are commonly known as baccharises but sometimes referred to as "brooms", because many members have small thin leaves resembling the true brooms. They are n ...

'' species are nectar sources for most of the predatory wasps, native skippers (small butterflies), and native flies in their ranges. ;Subspecies *''Baccharis pilularis'' subsp. ''consanguinea''(DC.) C.B.Wolf — primarily in coastal chaparral *''Baccharis pilularis'' subsp. ''pilularis'' — sandy coastal bluffs and beaches in California.

Cultivation

''Baccharis pilularis'' is cultivated as an

ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that i ...

, and used frequently in

drought tolerant Drought tolerance is the ability to which a plant maintains its biomass production during arid or drought conditions. Some plants are naturally adapted to dry conditions'','' surviving with protection mechanisms such as desiccation tolerance, detox ...

native plant In biogeography, a native species is indigenous to a given region or ecosystem if its presence in that region is the result of only local natural evolution (though often popularised as "with no human intervention") during history. The term is equ ...

, and

wildlife garden A wildlife garden (or wild garden) is an environment created by a gardener that serves as a sustainable haven for surrounding wildlife. Wildlife gardens contain a variety of habitats that cater to native and local plants, birds, amphibians, rep ...

s, and in

natural landscaping Natural landscaping, also called native gardening, is the use of native plants and adapted species, including trees, shrubs, groundcover, and grasses which are local to the geographic area of the garden. Benefits Maintenance Natural landsc ...

and

habitat restoration Restoration ecology is the scientific study supporting the practice of ecological restoration, which is the practice of renewing and restoring degraded, damaged, or destroyed ecosystems and habitats in the environment by active human interrupt ...

projects. The

cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...

ground cover Groundcover or ground cover is any plant that grows over an area of ground. Groundcover provides protection of the topsoil from erosion and drought. In an ecosystem, the ground cover forms the layer of vegetation below the shrub layer known as t ...

selections have various qualities of height and spread, leaf colors, and textures. The upright forms are useful for hedges and fence lines, and year-round foliage. Coyote brush is usually deer-resistant. The plants are also drought tolerant after maturity, requiring watering once a week until established, and then about once per month during the first summer. They can mature in one to two years. The plants prefer good drainage. Only male plants of ''Baccharis pilularis'' are cultivated for landscaping use. If these are substituted for ''Baccharis pilularis'' subsp. ''consanguinea'' in ecological restoration, there will not be as much seed set, nor recruitment of new individuals.

Cultivars

Cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture, ...

s, often with the common name "dwarf coyote brush" or "dwarf baccharis" indicating

selections, include: * ''Baccharis pilularis'' 'Pigeon Point'—from Pigeon Point, California coast. * ''Baccharis pilularis'' 'Twin Peaks'—from coast along Sonoma to Monterey Counties. * ''Baccharis pilularis'' 'Santa Ana'Las Pilitas: ''Baccharis pilularis'' ' Santa Ana'
/ref>

References